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After Niles runs off with Daphne on the day of her intended wedding to Donny, Mel is humiliated and bitter.
She pays Niles back by refusing to grant an immediate divorce; instead, she forces Niles to play along that the two remain happily married, assuring him that after he has done this on several select social occasions she will file for divorce.
She doesn't want anyone to know that she and Frasier are consummating their relationship and asks him not to tell.
He is covert with friends and attempts discretion with his family members, but in both cases it is interpreted as loneliness.
He states he is tired of the pity and ends up stating that he met a supermodel in Acapulco who would like him to be discreet.
Mel also has one son, and it is specifically stated she is divorced.
They don't believe him until nearly the end of the episode when they finally do see the two of them together.
Veronica "Ronee" Lawrence Crane (Wendie Malick) is Niles and Frasier's old babysitter, and the first woman to break Frasier's heart (he used to watch her kiss her boyfriend through the balusters).
The series ends with Frasier on a plane, but it is revealed at the last minute that he had chosen love over career as the plane lands in Chicago, where he hopes to reunite with Charlotte.
Kelly Easterbrook (Sela Ward) is a supermodel, part-time, in order to support her zoology Ph D work.